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Yes, I know this is not new in any way - just not firmly established so that we all use the same terminology, but that's the purpose of our project, right? For now I and many just avoid the problem by using informal terms like anterior and posterior fork. By the way, Drepanocercus spinistylus was moved to Acomoptera by Kerr (2011) and these species show a considerable intraspecific variation in the length and attachment of the "posterior" fork.

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---Original Message-----
From: dsamorim [mailto:dsamorim@usp.br]
Sent: 26. august 2014 13:24
To: mycetophylo@sciaroidea.info
Subject: Re: Thanks, Jostein! The most

There is a photo of exactly the same part of the wing of Paratinia sciarosoma in Amorim & Rindal 2007, Fig. 173. It actually suggests that the " cubital veins forking distally" is a secondary achievement of most Mycetophilidae due to a fusion of the basal parts of M4 and CuA. The long "cubital fork", with m-cu forking from CuA close to the wing base and connected to M1+2 is a condition seen in Palaeognoriste , Rhynchoheterotricha , Bolitophila , Chiletricha , Heterotricha , etc. Only some few mycetophilids preserve a pretty plesiomorphic condition, e.g., Drepanocercus .

----- Mensagem original -----

> De: "Jostein Kjaerandsen"
> Para: mycetophylo@sciaroidea.info
> Enviadas: Terça-feira, 26 de Agosto de 2014 7:04:07 > Assunto: RE: Thanks, Jostein! The most

> Files:
> http://sciaroidea.info/sites/sciaroidea.infosciaroidea.info/files/m-cu-W...

> Anyhow I think we need to settle on a new Cu terminology. Check out > these > Mycomya, Paratinia and Acomoptera wings - should make it clear that > the > Cu-fork concept is wrong.

> Cheers
> Jostein

> -----Original Message-----
> From: vblago [mailto:vblago@gmail.com] > Sent: 26. august 2014 00:16 > To: mycetophylo@sciaroidea.info > Subject: Re: Thanks, Jostein! The most

> On 25 August 2014 22:44, dsamorim wrote:

> > not sure if this will help or make it even more > complicated >

> That's why I am not sure if we have to get into this. For the purpose > of our > project it is enough to establish homology whithin Sciaroidea and > Bibionomorpha, which is rather straightforward. But let's see what > you > student will find. I'll check what material we have in NHM

> Cheers,
> Vlad

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