Species Page

Leptomorphus mandelai

Etymology: 
This species is named in honour of former South African President Nelson R. Mandela, in recognition of his role in ending apartheid in South Africa and for his advocacy of peace, reconciliation and social justice.
Habitus: 
The only extant species of Leptomorphus with the following combination of characters: laterotergite, anepisternum and anepimeron yellow; scutellum yellow; scutum mostly dark brown with yellow restricted to anterior and lateral margins; abdomen with tergite 7 brown (Fig. 23); male genitalia with tergal evagination bearing ventrally-directed fold along most of posterior margin and small point on apicomedial corner (Fig. 109). This species can be distinguished from other Afrotropical species by the yellow scutellum, brown tergite 7 (Fig. 23), and lack of small setae covering the scutum (except along dorsocentral lines).
Distribution: Text: 
South Africa: KwaZulu-Natal; Howick district, Karkloof Range, Geekie’s Farm (28.16°S, 30.21°E)

Macrorrhyncha anatolica

Type of Type: 
Holotype
Type Specimens Data: 
Male, Turkey, Aydın, Çine, Eskiçine Village, Çine Stream, 37°31’ N/ 28°04’ E, 80 m a.s.l., 21.04.2005, leg. H. KOÇ, A. KARAMAN&O. OZGÜL (in HNHM)
Distribution: Text: 
Turkey

Hemisphaeronotus

Type Discussion: 
Type species. Hemisphaeronotus rotundatus SAIGUSA, 2007
Etymology: 
The generic name is based on the hemispherically humpbacked mesoscutum; hemisphaera (hemisphere) + notus (back). Gender masculine.
Head: Shape: 
Head moderately large, hemispherical, in dried specimens tightly attached to more or less concaved cervical and prothoracic regions under anterior 1/3 of mesoscutum, therefore head directed ventrally and morphological posterior margin of head almost dorsal. Frons 1.3 times as wide as long, more or less narrowed anteriorly; face twice as wide as long; clypeus as long as face, twice as long as wide; frons, face and clypeus without setae. Vertex short, greater part of vertex and occiput representing bottom surface of hemispherical head concealed in lateral aspect, short setose only on upper lateral areas of vertex.
Head: Ocelli: 
Ocelli 3 in number, arranged almost transversely, but middle ocellus slightly shifted anteriorly from lateral pair, middle ocellus half as large as lateral pair in diameter, lateral pair situated midway between middle ocellus and inner eye margin.
Head: Eyes: 
Compound eyes oval, with inner margin almost straight, not emarginated above antennal bases, densely covered with inter-mmatidial setae 3 times as long as diameter of ommatidium; vertical diameter of eye 1.4 times as long as transverse diameter.
Head: Antennae: 
Antenna moderately long for Grioristini, longer in male than in female; scape as long as thick and short setose, pedicel spherical , half as long as scape, and short setose on ventral surface; flagellum consisting of 14 flagellomeres , each flagellomere pale pilose and bearing several short black setulae more abundant on basal flagellomeres; 1st flagellomere several times as long as thick; distal flagellomere almost as long as penultimate.
Head: Mouthparts: 
Mouthparts short; labrum triangular, almost as long as face; labella small , very short setose; maxillary palpus greatly shortened, 2/3 as long as height of compound eye, 1st and 2nd palpomeres united into short segment with some setulae, 3rd spherical and with large sensory area occupying most of mesal surface and bearing some 10 setulae, 4th cylindrical , as long as 3rd , 5th longer than 4th and with several setulae towards tip
Sizes: 
Medium- to large-sized
Habitus: 
Elongate, long-legged mycetophilids with strongly humpbacked mesoscutum overlapping head, incompletely divided mesepimeron, only short setose mesoscutellum lacking marginal setae, wing characters as in genus Boletina Staeger, 1840, male epandrium completely fused with gonocoxites bearing trifurcate gonostylus, and female cercus lobate and one-segmented.
Thorax: Notum: 
Mesoscutum large, hemispherical and strongly humpbacked,expanded anteriorly like an eave that overlaps head; parapsidal suture nearly transverse,directed medially at right angle to lateral margin of mesonotum. Acrostichals absent, dorsocentrals uniserial to irregularly biserial; sublateral portions of mesoscutum setose, prealar (notopleural) to supra-alar areas close to lateral margins of mesoscutum densely clothed with long stiff setae arranged in uniserial or irregularly biserial rows. Scutellum moderately large, oval and about 1.5 times as wide as long, bearing only short setulae on lateral areas of anterior half, setulae may be reduced to 1-2 in number and confined to lateral corners of scutellum.
Thorax: Lateral Sclerites: 
Mediotergite large and elongate , bare and not swollen posteriorly. Prothoracic and cervical areas situated on sunken ventral surface of mesoscutum; anterior pronotum with median area invisible in dried condition, with lateral part flattened bearing several setae; proepisternum bare, proepimeron narrowly tapered to sharpl y pointed ventral extremity that is articulated with anterior margin of mesokatepisternum ; precoxal sclerotization of prothorax bare , isolated as sclerite on membranous area; prosternum bare. Mesothoracic pleuron deep and much obliquely situated on lateral part of thorax, its vertical length 0.6-0.7 times as long as combined length of mesoscutum and mesoscutellum; no setae on any part of mesothoracic pleuron ; pleural suture of mesothorax appearing along only dorsal half of episternum and close to base of cx2; anapleural suture disappearing on anterior 2/3 to 3/4 of its ordinary length, therefore suture not reaching anterior margin of episternum; laterotergite large and bare. Metathoracic pleuron small, metakatepisternum twice as long as deep, quadrate and bare; metepimeron small and bare; metanotal seta just dorsal to halter base present or absent.
Thorax: Halters: 
Halter brown or yellow, with yellow shaft.
Wings: Sizes and Ratios: 
Wing elongate
Wings: Venation: 
Sc, R, R1, R4+5, r-m crossvein bearing long setae on dorsal surface, R, R1, apical part of R4+5 with longish setae on ventral surface in some species; M1, M2, M4 and apical part of CuA and CuP with short setae on dorsal surface. C slightly extended beyond tip of R4+5, Sc ending at or slightly proximad of Rs base, sc-r and R4 absent, r-m crossvein twice as long as Rs and slightly longer than common stem of M1 and M2: fork of M4 and CuA between level of r-m crossvein and fork of M1 and M2: cupseudovein (traditional CuP) distinct, CuP (traditional A1) strong, both extending near level of fork of M4 and CuA. Wing weakly infuscate on apical 1/4, with weak cloud from Rs to fork of MI and M2.
Wings: Setation: 
Wing membrane densely covered with microtrichia, lacking macrotrichia.
Legs: Sizes and Ratios: 
Legs long and slender as in other Gnoristini genera
Legs: Setation and Ornamentation: 
cx1 and cx2 setose on anterior surface, cx3 setose on outer surface; distal setae strong; femora short setose, with anteroventral row of setae as long as or half as long as f thickness ; tibiae with several anterior, dorsal, posterior and ventral short bristles, more or less thickened at apical portion; tl with oblong anterodistal patch densely clothed with minute pile; tarsi short haired bearing a few setae; apex of 5th tarsomere produced into dorsal process. Ungues bearing several ventral teeth, larger apically; in males of some species posterior unguis lamellate, with small apical projection; empodium small.
Abdomen: Colour: 
Abdomen elongate and moderately long setose
Abdomen: Segments: 
1st abdominal sternum entire, undivided into main anterior part and small posterior part, lacking setae; other abdominal sternum complete, without longitudinal fold lines. In male, 7th and 8th abdominal segments usually almost concealed in 6th segment; 7th segment 0.5-0.7 times as long as 6th, 7th tergum very short, represented by transverse linear sclerite lacking setae, 7th sternum setose; 8th segment shorter than 7th, 8th tergum similar to 7th tergum: 8th sternum bare or with a few setulae along posterior margin
Male Terminalia: Size and General Shape: 
Male genitalia large, wider than apical abdominal segments; epandrium fused with dorsal margins of gonocoxite leaving no border between them, more or less produced posteriorly like an eave and bearing some stiff setulae on distal margin
Male Terminalia: Gonocoxites: 
Gonocoxite with ventrodistal part produced in lamellate projection more or less triangular in shape
Male Terminalia: Gonostili: 
Gonostylus trilobate, dorsal lobe style-like, middle main lobe large and usually spatulate, densely clothed with minute spinules on flexed surface apically, ventral lobe small and setose
Male Terminalia: 10th tergite and cerci: 
Cercus small, membranous, concealed below eave-like expansion of epandrium and only short setose; pair of hypoproct lobes similar to cercus in shape and more sparsely setose; phallic organ small , compact, outwardly short produced into a stout projection with a minute tooth-like tip apically , and pair of short internal apophyses.
Female Terminalia: Tergites and Cerci: 
10th tergum (epiproct) consisting of lateral setiferous lobe and transversely narrow median part raised dorsally and roundly produced posteriorly
Female Terminalia: Colour, Size and General Shape: 
Female terminalia short; 7th abdominal segment 2/3 as long as 6th segment, 8th to 10th terga very short, 8th sternum with a pair of small lamellate lobes bearing short setae; 9th tergum nearly half as long as 8th, lacking setae, 9th sternum a long triangular plate applied just beneath hypoproct (10th venter)
Female Terminalia: Sternites and Gonopodes: 
10th sternum (hypoproct) divided into pair of elongate sclerites arising from epiproct without distinct border and extending posteriorly below cerci and bearing some setae; cercus unsegmented, comprising longish, broadly lamellate, oval lobe , short setose and bearing some longish setae along outer margin.

Diadicidia ferruginosa

Head: Antennae: 
Third flagellomere, length-width: m = 2.3 to 3.2, f = 2.69 to 3.2. Fourth flagellomere: m = 1.64 to 2.22, f = 2.16 to 2.50. Report lengths 3rd article/4e: m = 1.2 to 1.8, f = 1.21 to 1.83. Basal third or half of first flagellomere yellowish-brown, the rest of the antenna dark brown.
Habitus: 
Coloration variable; body generally dark brown, but sometimes almost entirely yellow. Mesonotum sometimes with with three separate brown bands. Scutellum yellow to yellow-brown. Legs yellow, tibiae and tarsi brown.
Wings: General: 
wings slightly smoky gray
Wings: Sizes and Ratios: 
Length of the wing: M = 3.3 mm to 3.7 mm, f = 4.0 to 4.7 mm.
Wings: Venation: 
R1 ending above the base M-fork; Sc ending before the base RS; RS distally slightly bent.
Male Terminalia: 9th tergite: 
Tergite IX triangular, slightly rounded at apex, as long as its width, with numerous clear setae
Male Terminalia: Gonostili: 
Apex of gonostilus straight, slender, bifurcate
Male Terminalia: Aedeagus and Apodemes: 
Aedeagus relatively narrow, only slightly expanded distally

Palaeodocidia ishizakii

Distribution: Text: 
Japan

Diadocidia (Diadocidia) halopensis Sevcik, 2003

Type of Type: 
Holotype
Type Specimens Data: 
Male; Papua New Guinea, Madang Province, Halopa village, 600 m, primary rain forest, November 2000 (L. Cizek leg., Malaise trap): [Bishop Museum, Honolulu]
Etymology: 
After Halopa village
Head: Coloration: 
dark brown
Head: Ocelli: 
3, lateral ocelli about twice as large as median, and separated from the eye margin for a distance about 1.5 times their diameter
Sizes: 
body length 2.4 mm
Habitus: 
general coloration brown, legs yellowish brown
Thorax: Halters: 
brown
Wings: Sizes and Ratios: 
length 2.0 mm, length to width ratio 2.3
Wings: Setation: 
covered with macrotrichia, without microtrichia

Diadocidia (Adidocidia) papua Sevcik, 2003

Synonyms: 
none
Type of Type: 
Holotype
Type Specimens Data: 
Male, Papua New Guinea, Morobe District, Mt. Missim, 2350m, 20-31.10.1966 (G.A.Samuelson): [BMH]
Etymology: 
Refers to Papua New Guinea, singular nominative noun.
Head: Coloration: 
Yellowish brown, antennae brown
Head: Antennae: 
Scape and pedicel short, as long as wide, covered with setae. Flagellum cylindrical, densely covered with fine setulae, tapering toward apex, with 14 flagellomeres. Ratio of length to width for the first flagellomere is 4.5, for the flagellomeres 2 to 8 is 2.3.
Head: Mouthparts: 
Clypeus rounded and setose. Mouthparts and palpi yellowish brown. Palpi with 4 palpomeres, covered with dark setae. Palpomere 4 (apical) narrow and somewhat darkened. Relative length of palpomeres 1 to 4: 1:2:3.2:3.6.
Sizes: 
Body Sizes: 4.3 mm, antenneae 3.2 mm, flagellum 3.1 mm Wings: 4.6 mm
Habitus: 
General coloration dark brown, legs yellowish
Thorax: Notum: 
Mesonotum yellowish brown, with three longitudinal dark stripes, which are covered with dark setae. Lateral margins of mesonotum also darker and setose. Scutellum brown, with several long dark apical bristles. Mediotergite bare, yellowish brown
Thorax: Lateral Sclerites: 
Laterotergites and preepisternum 2 bare, yellowish brown. Anepisternum with about 10 black setae near upper margin. Antepronotum with 4 black setae.
Thorax: Halters: 
Yellowish, covered with dark setulae.
Wings: Sizes and Ratios: 
Ratio of wing length to width 2.6. Ratio of the length of R1 to the total wing length is 0.67. M ratios: 0.54 and 0.71. Cu ratios: 0.63 and 1.38. A1 reaching wing margin.
Wings: Venation: 
Costa produced beyond R5. Sc ending in C before base of Cu-fork. Sc2 absent. R1 ending in C well beyond base of M-fork. Veins R-M and bM-Cu in one line.
Wings: Setation: 
A1 setose
Wing: Basalia: 
Calypter well developed, bearing long setae along its margin.
Legs: Sizes and Ratios: 
Femora laterally compressed and thickened medially. Fore and mid femora somewhat longer than coxae, hind femur almost twice as long as hind coxa. Ratio of the spur length to the length of the first tarsomere for legs are: 0.23; 0.26; 0.31. All spurs on mid and hind tibiae subequal in length. Ratio of femur (including trochanter) to tibia for legs: 0.83; 0.86, 0.75. Ratio of tibia to tarsus: 0.62; 0.87; 1.17. Ratio of first tarsomere to tarsus: 0.50; 0.52; 0.52.
Legs: Setation and Ornamentation: 
Legs covered with dark trichia and setae. All coxae with a longitudinal row of black setae, c1 anterolaterally, c2 laterally and c3 posterolaterally. Femora clothed with numerous trichia and with a row of longer setae ventrally.
Legs: Colour: 
Yellowish
Abdomen: Colour: 
Brown
Abdomen: Segments: 
All tergites and sternites covered with dark setae.
Male Terminalia: Colour: 
Brown, apical part of gonostili and aedeagus black.
Male Terminalia: Size and General Shape: 
Length 0.5 mm.
Male Terminalia: 9th tergite: 
Semicircular, almost as long as wide, densely covered with long setae
Male Terminalia: Gonocoxites: 
Caudal margin of gonocoxites ventrally with wide V-shaped depression.
Male Terminalia: Gonostili: 
Long and narrow, with two black apical branches. Between the branches is a comb-like structure.
Distribution: Text: 
Papua New Guinea
Hosts and Substrates: 
unknown

Diadocidia fissa Zaitzev, 1994

Type Specimens Data: 
Male; Russia: Kostroma Region, Ugory, 7.09.1981 (A.Zaitzev) [ZIN]
Head: Coloration: 
Dark brown
Head: Antennae: 
Two-coloured, scape, pedicell and ventral part of the first plagellomere yellow, the rest of flagellomeres dark brown. Length of middle flagellomeres only slightly more than its width
Head: Mouthparts: 
Yellow; palpi dark fulvous. Basal palpomere longer than the second palpomere
Sizes: 
Wing 4.5 mm
Thorax: General: 
Yellow-brownish, mosonotum yellowish, with indistinct fulvous stripes, pleurae yellow, scutellum and mediotergite yellow
Thorax: Lateral Sclerites: 
Lateral sclerites: Anepisternum with a group of small setae
Distribution: Text: 
Russia: Kostroma

Diadocidia furnacea Chandler, 1994

Type Specimens Data: 
Male; Israel: Monfort, 14.03.1985 (A.Freiberg) [TAU]
Etymology: 
Name refers to the darker brown body coloration of the species as compared to the other Palaearctic species
Head: Coloration: 
Brown, grey dusted, antenna dark brown
Head: Ocelli: 
Lateral ocelli close to eye margins
Head: Eyes: 
Eyes not produced dorsally
Head: Antennae: 
With long setae on short basal segments; flagellum long, slender, with segments 3 times as long as broad
Sizes: 
Wing 4.0 mm
Thorax: Notum: 
Shining dark brown with grey dusting on acrostichal and dorsocentral rows, which are in 1 or 2 series, with bare stripes between these series and between the dorsostichal and the long lateral bristling; scutellum with 2 pairs of long setae among series of short marginals.
Thorax: Lateral Sclerites: 
Pleura and mediotergite bare
Thorax: Halters: 
Brownish yellow
Wings: General: 
Yellowish with brown veins
Wings: Venation: 
Sc ends on C; Sc2 before middle of vein Sc and well before base of vein RS; vein R1ends at nearly two thirds of wing length; vein R4 absent; vein R5 downturned apically; costa extends 0.4 distance from tip of vein R5 to that of vein M1; crossvein r-m, vein tb and base of vein CuA1 (M3+4) form irregular line just beyond basal third of wing, vein CuP end near the base of posterior fork; vein A1 reaches wing margin.
Wings: Setation: 
Mostly covered with both macrotrichia and microtrichia.
Legs: Setation and Ornamentation: 
With long dark setae on coxae and short irregular setae on all parts; mid tibia with 2 short posterior setae; hind tibia with series of short strong setae: 2 anterior, 3 dorsal and 4 posterior on apical half; tibial spurs more than twice apical tibial width
Legs: Colour: 
Yellow
Male Terminalia: Size and General Shape: 
Closely resembles D. ferruginosa
Male Terminalia: 9th tergite: 
More rounded apical margin than in D. ferruginosa
Distribution: Text: 
Israel

Diadocidia (Adidocidia) trispinosa Polevoi, 1996

Taxonomic Discussion: 
Close to D. borealis, but has three strong apical setae on gonostili instead of one. The holotype specimen was misidentified as D. ferruginosa in Polevoi, 1995
Type Specimens Data: 
Male; Finland: Pihru, about 200 m above see level; 4-5.08.1993 (A.Polevoi, malaise trap) [ZIN]
Head: Coloration: 
Black, mouthparts and palpi brown, antennae black, scape and pedicell brown
Head: Antennae: 
Sixth flagellomere about 3 times as long as wide
Sizes: 
Wing 3.6 mm
Thorax: Notum: 
Dark brown, grey dusted with lighter humeral spots
Thorax: Lateral Sclerites: 
Pleurae dark brown, propleurae somewhat lighter
Wings: General: 
Greyish
Legs: Sizes and Ratios: 
t1:bt1 = 1.1; t2:bt2 = 1.6; t3:bt3 = 2.2
Legs: Colour: 
Yellowish brown
Abdomen: Colour: 
Dark brown
Male Terminalia: Colour: 
Brown
Male Terminalia: 9th tergite: 
With apical bristles not thickened
Male Terminalia: Gonostili: 
With two apical teeth and three strong setae
Distribution: Text: 
Finland
Scratchpads developed and conceived by: Vince Smith, Simon Rycroft, Dave Roberts, Ben Scott...