Phylogeny of the tribe Sciophilini (Diptera: Mycetophilidae: Sciophilinae)

Publication Type:Journal Article
Year of Publication:2013
Authors:Borkent, CJ, Wheeler, TA
Journal:Systematic Entomology
Volume:38
Pagination:407-427
ISSN:1365-3113
Abstract:

The monophyly and phylogenetic relationships within the species rich Sciophilini (Diptera: Mycetophilidae) were analysed, based on 96 adult morphological characters. The cladistic analysis included 80 Sciophilini exemplar species (representing all but 1 of the 36 genera placed previously in the Sciophilini) and 11 outgroup taxa of other mycetophilid tribes. The monophyly of Sciophilini was supported in the parsimony analysis by four synapomorphies. The tribe now contains 34 genera: Acnemia Winnertz, Acomoptera Vockeroth, Adicroneura Vockeroth, Afrocnemia Matile, Allocotocera Mik, Anaclileia Meunier, Aneura Marshall, Austrosciophila Tonnoir, Azana Walker, Baeopterogyna Vockeroth, Cluzobra Edwards, Drepanocercus Vockeroth, Duretophragma Borkent gen.n., Eudicrana Loew, Leptomorphus Curtis, Loicia Vockeroth, Megalopelma Enderlein, Monoclona Mik, Morganiella Tonnoir & Edwards, Neoallocotocera Tonnoir, Neoaphelomera Miller, Neotrizygia Tonnoir & Edwards, Neuratelia Rondani, Paramorganiella Tonnoir, Paratinia Mik, Paratrizygia Tonnoir, Parvicellula Marshall, Phthinia Winnertz, Polylepta Winnertz, Sciophila Meigen, Stenophragma Skuse, Tasmanina Tonnoir, Taxicnemis Tonnoir & Edwards, and Trizygia Skuse. Four genera placed previously in Sciophilini (Coelophthinia Edwards, Impleta Plassmann, Speolepta Edwards and Syntemna Winnertz) are transferred to the Gnoristini. Neoneurotelia Shinji and Neoparatinia Shinji are considered nomina dubia. Diagnoses are given for all genera in the tribe. Duretophragmagen.n. is described for the following species (all of which are comb.n.): Duretophragma andina (Duret), Duretophragma argentina (Duret), Duretophragma glabanum (Johannsen), Duretophragma fusca (Edwards), Duretophragma humeralis (Edwards), Duretophragma intermedia (Edwards), Duretophragma longifurcata (Freeman) (type species), Duretophragma morigenea (Edwards), Duretophragma naumanni (Duret), Duretophragma nigricauda (Edwards), Duretophragma obscura (Duret), Duretophragma ochracea (Freeman), Duretophragma pleuralis (Edwards) and Duretophragma similis (Johannsen). Other new generic combinations include: Trizygia albidens (Oliveira & Amorim) comb.n., Trizygia alvesi (Oliveira & Amorim) comb.n., Trizygia balbi (Oliveira & Amorim) comb.n., Trizygia camargoi (Oliveira & Amorim) comb.n. and Afrocnemia stellamicans (Chandler) comb.n.

URL:http://dx.doi.org/10.1111/syen.12002
DOI:10.1111/syen.12002
Taxonomic name: 
Gnoristinae (Classification), Acomoptera (Classification), Adicroneura (Classification), Coelophthinia (Classification), Drepanocercus (Classification), Impleta (Classification), Speolepta (Classification), Syntemna (Classification), Sciophilinae (Classification), Acnemia (Classification), Afrocnemia (Classification), Allocotocera (Classification), Anaclileia (Classification), Aneura (Classification), Austrosciophila (Classification), Azana (Classification), Baeopterogyna (Classification), Cluzobra (Classification), Duretophragma (Classification), Duretophragma andina (Classification), Duretophragma argentina (Classification), Duretophragma glabanum (Classification), Duretophragma humeralis (Classification), Duretophragma intermedia (Classification), Duretophragma longifurcata (Classification), Duretophragma morigenea (Classification), Duretophragma naumanni (Classification), Duretophragma nigricauda (Classification), Duretophragma obscura (Classification), Duretophragma ochracea (Classification), Duretophragma pleuralis (Classification), Duretophragma similis (Classification), Eudicrana (Classification), Leptomorphus (Classification), Loicia (Classification), Megalopelma (Classification), Monoclona (Classification), Morganiella (Classification), Neoallocotocera (Classification), Neoaphelomera (Classification), Neoneurotelia (Classification), Neoparatinia (Classification), Neotrizygia (Classification), Neuratelia (Classification), Paramorganiella (Classification), Paratinia (Classification), Paratrizygia (Classification), Parvicellula (Classification), Phthinia (Classification), Polylepta (Classification), Sciophila (Classification), Stenophragma (Classification), Tasmanina (Classification), Taxicnemis (Classification), Trizygia (Classification)
Groups audience: 
Taxonomy checked: 
Yes
TDWG distribution: 
No
Specimens imported: 
No
Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith